The mammalian eye is a complex organ that tranduces light into patterns of nerve impulses. The transduction takes place
in the retina, a layer of photosensitive cells at the back of the eye. It contains two types of receptors called rods and cones
Rods and cones are classified as secondary recpetors - they are
more complex than primary receptors like pacinian corpuscle. A
primary receptor consists of a single neurone. In secondary
receptors, a modified epithetial cells detect changes in the
stimulus and passes this information onto a seperate neurone
which transmits the information as nerve impulses
Rods and cones are part of a sense
organ, the eye. Sense organs are
complex stimulus-gathering structures
consisting of group sensory receptors
Up to 150 rods may connect via synapses to one
neurone, a characteristic known as convergence.
This enables a group of rods to function as a
photosensitive unit, gathering light from a relatively
wide area and combining its stimulatory effects.
Convergence enables rods to provide photosensory
information at low light intensities, but this increased
sensitivity is at the expence of visual acuity (the
ability to discriminate fine detail)
Consequently, in dim light when cones are not
functioning, the rods provide only enough
information for an ill-defined informationthat lacks
colour. Also because rods are the only type of
photoreceptor at the outmost edge of the retina,
something 'out the corner of your eye' lacks detail
A rod cell has in its outer segement up to 1000 vesicles, each
containing a photosenitive pigment called rhodopsin. Rhodopsin is
made up of the protein opsin and retinal, a derivative of vitamin A.
Light causes retinal to change shape, which results in retinal and
opsin breaking apart.; a process called bleaching. This triggers a
series of events which alters the permeability of the rods plasma
membrane and contributes to the fromation of a generator
potential in the sensory neurone. The generator potential results
from the combined effects of all rods serving a particuar sensory
neurone. When it exceeds a critical threshold level, a nerve
impulse is transmitted.
Cones enable us to see colour and fine details. There are three types of
cone, each containing a different form of thr pigment iodopsin which breaks
down only bright light. The three pigments together are sensitive over the
visible spectrum, but on is most sensitive to blue light, one to green light, and
the other to red light. According to the trichromatic theory, different colours
are percieved by mixing the information from the different types of cone.
Whereas rods function in groups connected to one sensory neurone, a
single cone may have its own sesonry neurone. if light of suffieciently
high intensity from two seperate sources falls on two such cone cells,
two seperate impulses are transmitted along two sensory neurones and
the brain recieves two seperate images. Thus cones provide much
more detaild sensory information, but only at high light intensities
The fovea and blind spot. Over most of
the retina, rods and cones are buried
under a layer of blood vessels and
nerve fibres which lead into the optic
nerve. However, the fovea, a small
depression in the retine opposite the
lens, consists only of cones. The layer
of nerve fibres here is thin and there are
no cappilaries. This means that light
falling on the fovea produces a clear,
well defined visual image in colour.
When a person wants to examine the
fine details of an object, they eyes move
automatically so that the light from the
object falls on the fovea.
The eyes adapt to different
levels of brightness by varying
theor sesnitivity. A prelonged
period in a darkened room
results in photosensitive
pigments being formed much
faster than they are being
broken down, thus increasing
the sesitivity to light. This
process is called dark
adaptation. A prelonged period
in bight light results in
photosenstive pigments in rods
and cones being broken down,
reducing sensitivity to light. This
is called light adapatiation